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African American Percy Julian was a scientist and entrepreneur whose work helped people around the world to see. Named in 1999 as one of the greatest achievements by a US chemist in the past hundred years, ________ led to the first mass-produced treatment for glaucoma.

Which choice completes the text so that it conforms to the conventions of Standard English?
Difficulty: Medium

Julian synthesized the alkaloid physostigmine in 1935; it


in 1935 Julian synthesized the alkaloid physostigmine, which


Julian's 1935 synthesis of the alkaloid physostigmine


the alkaloid physostigmine was synthesized by Julian in 1935 and

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