A pioneer in the field of taphonomy (the study of how organic remains become fossils), ________ may be just as prevalent in the fossil record as those of thick-shelled organisms.

Which choice completes the text so that it conforms to the conventions of Standard English?

Question

Accepted Answer

Anna Behrensmeyer demonstrated in a 2005 analysis that fossils of thin-shelled organisms

Answer

fossils of thin-shelled organisms, Anna Behrensmeyer demonstrated in a 2005 analysis,

Answer

it was demonstrated in a 2005 analysis by Anna Behrensmeyer that fossils of thin-shelled organisms

Answer

a 2005 analysis--by Anna Behrensmeyer--demonstrated that fossils of thin-shelled organisms